Hi bccc2005,
The two black ink tanks serve different purposes and are used for different types of printing, so one would not be used in place of the other, should one of the inks run out. The larger black ink tank is a pigment-based ink and is used for your document printing when plain paper is selected, you are doing single-sided printing, and your document contains black text. The smaller ink tank is a dye-based ink and is used when printing photos, performing duplex (double-sided) printing, and when a paper type other than plain paper is selected (i.e., envelopes, labels, photo paper, etc.).
Hope this helps!
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